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| + | Smallmouth bass have been sporadically captured below Glen Canyon Dam and in Grand Canyon since 2003. They were first introduced into Lake Powell in 1982 [https://wayneswords.net/threads/smallmouth-bass-history-in-lake-powell.1340/] and there are populations in ponds and lakes in the upper Little Colorado River as well as in Lake Mead. |
| + | |
| + | '''2023''' <br> |
| + | Annual catch of smallmouth bass through sampling by USGS and AZGFD in the Lees Ferry reach has averaged 0-3 fish per year until 2022. A fall 2022 USGS trip captured 30 young of year smallmouth bass (58-93 mm fork length (FL)), and 20+ young of year detected in the slough that year through sampling by NPS. This represents the first evidence of spawning below Glen Canyon Dam. Most smallmouth bass captured so far have come from the first 5 miles below Glen Canyon Dam (e.g., only three smallmouth bass were caught in the lower Lees Ferry site located around 4 mile bar, while 27 were captured just upriver of the slough) suggesting that the extent of the smallmouth bass invasion may still be limited to the upper portions of the river nearest Glen Canyon Dam. Evidence of the smallmouth bass invasion being limited to the Lees Ferry stretch is further supported by a September 2022 river-wide seining trip targeting backwaters for smallmouth bass which discovered zero smallmouth bass downstream of Lees Ferry. Further sampling by NPS and others in fall 2022 detected and removed over 300 young of year smallmouth bass in Lees Ferry.[http://gcdamp.com/images_gcdamp_com/0/06/2_Final_Technical_Report_and_Recommendation_Regarding_a_Spring_Flow_Experiment_at_Glen_Canyon_Dam%2C_April_2023_%281%29.pdf] |
| + | |
| + | '''2022''' <br> |
| PAGE, Ariz. —On July 1, 2022, juvenile smallmouth bass were found in the Colorado River below the Glen Canyon Dam. Smallmouth bass are a predatory non-native fish that have adversely impacted native and federally protected fish in the upper basin of the Colorado River over the last 20 years. The National Park Service (NPS) is working closely with partners and stakeholders to coordinate rapid response actions to address this finding in accordance with previously approved management plans. | | PAGE, Ariz. —On July 1, 2022, juvenile smallmouth bass were found in the Colorado River below the Glen Canyon Dam. Smallmouth bass are a predatory non-native fish that have adversely impacted native and federally protected fish in the upper basin of the Colorado River over the last 20 years. The National Park Service (NPS) is working closely with partners and stakeholders to coordinate rapid response actions to address this finding in accordance with previously approved management plans. |
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| The NPS encourages anglers to report any observations or catches of smallmouth bass below the dam. If caught, please remove the fish, put on ice and contact staff at Glen Canyon National Recreation Area or Grand Canyon National Park with a description of the location where found. Report findings to: Jeff Arnold, Glen Canyon National Recreation Area, e-mail us or Brian Healy, Grand Canyon National Park, e-mail us. [https://www.nps.gov/grca/learn/news/invasive-smallmouth-bass-colorado-river-below-glen-canyon-dam.htm] | | The NPS encourages anglers to report any observations or catches of smallmouth bass below the dam. If caught, please remove the fish, put on ice and contact staff at Glen Canyon National Recreation Area or Grand Canyon National Park with a description of the location where found. Report findings to: Jeff Arnold, Glen Canyon National Recreation Area, e-mail us or Brian Healy, Grand Canyon National Park, e-mail us. [https://www.nps.gov/grca/learn/news/invasive-smallmouth-bass-colorado-river-below-glen-canyon-dam.htm] |
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| + | The lower slough was treated with rotenone on September 17-19, 2022. |
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| + | |class="MainPageBG" style="width:45%; border:1px solid #cedff2; background:#f5faff; vertical-align:top;"| |
| + | {| width="100%" cellpadding="2" cellspacing="5" style="vertical-align:top; background:#f5faff;" |
| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Links</h2> |
| + | |- |
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| + | *[https://qcnr.usu.edu/coloradoriver/files/research/2024-SMB-Report-3.pdf Review of Smallmouth Bass Management in the Colorado River Ecosystem] |
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2023-02-16-amwg-meeting/20230216-InvasiveFishSpeciesBelowGlenCanyonDam-508-UCRO.pdf 2023 Invasive Fish Species Below Glen Canyon Dam: A Strategic Plan to Prevent, Detect and Respond] |
| + | *[http://gcdamp.com/index.php?title=National_Park_Service_Expanded_Non-native_Aquatic_Species_Management_Plan_and_EA 2019 National Park Service Expanded Non-native Aquatic Species Management Plan] |
| + | *[http://gcdamp.com/index.php?title=2013_National_Park_Service_Comprehensive_Fisheries_Management_Plan 2013 National Park Service Comprehensive Fisheries Management Plan] |
| + | *[http://gcdamp.com/index.php/Nonnative_Invasive_Aquatic_Species Nonnative Invasive Aquatic Species Page] |
| + | *[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf Habitat suitability information: Smallmouth bass] |
| + | *[https://gcdamp.com/index.php/The_-12_Mile_Slough The -12 Mile Slough Page] |
| + | |
| + | |- |
| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Projects</h2> |
| + | |- |
| + | |style="color:#000;"| |
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| + | *[https://coloradoriverrecovery.org/uc/wp-content/uploads/sites/2/2022/02/TechnicalReport-nna-Bestgen-2018-SmallmouthBassStudyPlan.pdf Evaluate effects of flow spikes to disrupt reproduction of smallmouth bass in the Green River downstream of Flaming Gorge Dam] |
| + | |
| + | |- |
| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Presentations and Papers</h2> |
| + | |- |
| + | |style="color:#000;"| |
| + | |
| + | '''2024''' |
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2024-02-29-amwg-meeting/20240229-FisheriesReviewAnnualReportingFY2023-508-UCRO.pdf Fisheries Review: Annual Reporting FY2023 ] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-01-25-twg-meeting/20240125-AnnualReportingMeeting-SmallmouthBassPopulationModelingImplications-508-UCRO.pdf Smallmouth Bass Population Modeling and Implications ] |
| + | |
| + | '''2023''' |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-TWGMeeting-SBAHGUpdateNonnativeFishStrategicPlan-508-UCRO.pdf SBAHG Update & Nonnative Fish Strategic Plan: Review and Discussion] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-AnnualReportingMeeting-SmallmouthBassDepletionEffortGlenCanyonNRA-508-UCRO.pdf Smallmouth Bass Depletion Effort in Glen Canyon National Recreation Area ] |
| + | |
| + | '''2022''' |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2022-10-13-twg-meeting/20221013-SmallmouthBassAdHocGroupReview-508-UCRO.pdf Smallmouth Bass Ad Hoc Group: Review, updates, and next steps ] |
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2022-05-18-amwg-meeting/20220518-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-508-UCRO.pdf Near-Term Threat of Smallmouth Bass Establishment below Glen Canyon Dam ] |
| + | *[https://www.usatoday.com/story/news/nation/2022/07/06/colorado-river-threatened-by-smallmouth-bass/7825588001/ 'Worst fears confirmed' in biologists' fight to save ancient Colorado River fish] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-FlowSpikeReduceSmallmouthBass-Presentation-508-UCRO.pdf Flow spike to reduce smallmouth bass reproductive success, Green River, 2021 ] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-Presentation-508-UCRO.pdf Near-Term Threat of Smallmouth Bass Establishment below Glen Canyon Dam ] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2022-01-13-twg-meeting/20220113-AnnualReportingMeeting-FishModellingSupportManagementDecisions-508-UCRO.pdf Fish modelling to support management decisions ] |
| + | |
| + | '''2016''' |
| + | *[https://www.researchgate.net/publication/315875119_RIVER_REGULATION_AFFECTS_REPRODUCTION_EARLY_GROWTH_AND_SUPPRESSION_STRATEGIES_FOR_INVASIVE_SMALLMOUTH_BASS_IN_THE_UPPER_COLORADO_RIVER_BASIN Bestgen and Hill. 2016. River regulation affects reproduction, early growth, and suppression strategies for invasive smallmouth bass in the upper Colorado River basin. ] |
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| Nest building and spawning occur when the water temperature is | | Nest building and spawning occur when the water temperature is |
| 12.8-21.0° C, but most activity occurs at or above 15° C. [https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] | | 12.8-21.0° C, but most activity occurs at or above 15° C. [https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] |
| + | |
| + | Mature females may contain 2000-15,000 golden yellow eggs. Males may spawn with several females on a single nest. On average each nest contains about 2,500 eggs, but nests may contain as many as 10,000 eggs. Eggs hatch in about 10 days if water temperatures are in the mid-50's (°F), but can hatch in 2-3 days if temperatures are in the mid-70's (°F). Males guard the nest from the time eggs are laid until fry begin to disperse, a period of up to a month. [https://www.fs.usda.gov/Internet/FSE_DOCUMENTS/fsbdev3_018493.pdf] |
| + | |
| + | Bass spawning videos |
| + | *[https://www.youtube.com/watch?v=50m8DoiKVVU Video 1] |
| + | *[https://www.youtube.com/watch?v=zBDDliPHdVk Video 2] |
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| '''Food: '''The diet of smallmouth bass changes from small to large food items as the | | '''Food: '''The diet of smallmouth bass changes from small to large food items as the |
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| populations of native fishes. [https://www.usbr.gov/lc/phoenix/biology/azfish/pdf/smallmouthbass.pdf] | | populations of native fishes. [https://www.usbr.gov/lc/phoenix/biology/azfish/pdf/smallmouthbass.pdf] |
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− | {| width="100%" cellpadding="2" cellspacing="5" style="vertical-align:top; background:#f5faff;"
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− | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Links</h2>
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| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Spike flows</h2> |
− | | + | |
− | *[http://gcdamp.com/index.php?title=Nonnative_Invasive_Aquatic_Species Nonnative Invasive Aquatic Species]
| + | |
− | *[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf Habitat suitability information: Smallmouth bass]
| + | |
− | *[http://gcdamp.com/index.php?title=National_Park_Service_Expanded_Non-native_Aquatic_Species_Management_Plan_and_EA 2019 National Park Service Expanded Non-native Aquatic Species Management Plan]
| + | |
− | *[http://gcdamp.com/index.php?title=2013_National_Park_Service_Comprehensive_Fisheries_Management_Plan 2013 National Park Service Comprehensive Fisheries Management Plan]
| + | |
− | | + | |
− | |-
| + | |
− | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Projects</h2>
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− | |style="color:#000;"|
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− | *[https://coloradoriverrecovery.org/uc/wp-content/uploads/sites/2/2022/02/TechnicalReport-nna-Bestgen-2018-SmallmouthBassStudyPlan.pdf Evaluate effects of flow spikes to disrupt reproduction of smallmouth bass in the Green River downstream of Flaming Gorge Dam]
| + | |
− | | + | |
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− | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Presentations and Papers</h2>
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− | '''2023'''
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-TWGMeeting-SBAHGUpdateNonnativeFishStrategicPlan-508-UCRO.pdf SBAHG Update & Nonnative Fish Strategic Plan: Review and Discussion]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-TWGMeeting-GlenCanyonDamSmallmouthBassEnvironmentalAssessmentUpdate-508-UCRO.pdf Glen Canyon Dam/Smallmouth Bass Environmental Assessment Update]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-AnnualReportingMeeting-SmallmouthBassDepletionEffortGlenCanyonNRA-508-UCRO.pdf Smallmouth Bass Depletion Effort in Glen Canyon National Recreation Area ]
| + | |
− | | + | |
− | '''2022'''
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2022-10-13-twg-meeting/20221013-SmallmouthBassAdHocGroupReview-508-UCRO.pdf Smallmouth Bass Ad Hoc Group: Review, updates, and next steps ]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/amwg/2022-05-18-amwg-meeting/20220518-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-508-UCRO.pdf Near-Term Threat of Smallmouth Bass Establishment below Glen Canyon Dam ]
| + | |
− | *[https://www.usatoday.com/story/news/nation/2022/07/06/colorado-river-threatened-by-smallmouth-bass/7825588001/ 'Worst fears confirmed' in biologists' fight to save ancient Colorado River fish]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-FlowSpikeReduceSmallmouthBass-Presentation-508-UCRO.pdf Flow spike to reduce smallmouth bass reproductive success, Green River, 2021 ]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-Presentation-508-UCRO.pdf Near-Term Threat of Smallmouth Bass Establishment below Glen Canyon Dam ]
| + | |
− | *[https://www.usbr.gov/uc/progact/amp/twg/2022-01-13-twg-meeting/20220113-AnnualReportingMeeting-FishModellingSupportManagementDecisions-508-UCRO.pdf Fish modelling to support management decisions ]
| + | |
− | | + | |
− | '''2016'''
| + | |
− | *[https://www.researchgate.net/publication/315875119_RIVER_REGULATION_AFFECTS_REPRODUCTION_EARLY_GROWTH_AND_SUPPRESSION_STRATEGIES_FOR_INVASIVE_SMALLMOUTH_BASS_IN_THE_UPPER_COLORADO_RIVER_BASIN Bestgen and Hill. 2016. River regulation affects reproduction, early growth, and suppression strategies for invasive smallmouth bass in the upper Colorado River basin. ]
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− | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Flow fluctuations</h2> | + | |
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| and Duever 1968).[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] | | and Duever 1968).[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] |
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− | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Other Stuff</h2>
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Updates
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Smallmouth bass have been sporadically captured below Glen Canyon Dam and in Grand Canyon since 2003. They were first introduced into Lake Powell in 1982 [2] and there are populations in ponds and lakes in the upper Little Colorado River as well as in Lake Mead.
2023
Annual catch of smallmouth bass through sampling by USGS and AZGFD in the Lees Ferry reach has averaged 0-3 fish per year until 2022. A fall 2022 USGS trip captured 30 young of year smallmouth bass (58-93 mm fork length (FL)), and 20+ young of year detected in the slough that year through sampling by NPS. This represents the first evidence of spawning below Glen Canyon Dam. Most smallmouth bass captured so far have come from the first 5 miles below Glen Canyon Dam (e.g., only three smallmouth bass were caught in the lower Lees Ferry site located around 4 mile bar, while 27 were captured just upriver of the slough) suggesting that the extent of the smallmouth bass invasion may still be limited to the upper portions of the river nearest Glen Canyon Dam. Evidence of the smallmouth bass invasion being limited to the Lees Ferry stretch is further supported by a September 2022 river-wide seining trip targeting backwaters for smallmouth bass which discovered zero smallmouth bass downstream of Lees Ferry. Further sampling by NPS and others in fall 2022 detected and removed over 300 young of year smallmouth bass in Lees Ferry.[3]
2022
PAGE, Ariz. —On July 1, 2022, juvenile smallmouth bass were found in the Colorado River below the Glen Canyon Dam. Smallmouth bass are a predatory non-native fish that have adversely impacted native and federally protected fish in the upper basin of the Colorado River over the last 20 years. The National Park Service (NPS) is working closely with partners and stakeholders to coordinate rapid response actions to address this finding in accordance with previously approved management plans.
Lower lake levels at Lake Powell and rising temperatures in the Colorado River have increased concerns that non-native, warm-water predators, including smallmouth bass, could pass through the dam and begin reproducing. If these warm-water predators establish populations below the dam, it is likely they would negatively impact native fish communities in Glen Canyon National Recreation Area and in Grand Canyon National Park.
The NPS encourages anglers to report any observations or catches of smallmouth bass below the dam. If caught, please remove the fish, put on ice and contact staff at Glen Canyon National Recreation Area or Grand Canyon National Park with a description of the location where found. Report findings to: Jeff Arnold, Glen Canyon National Recreation Area, e-mail us or Brian Healy, Grand Canyon National Park, e-mail us. [4]
The lower slough was treated with rotenone on September 17-19, 2022.
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Projects
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Presentations and Papers
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2024
2023
2022
2016
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Smallmouth bass overview
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Description: Smallmouth bass are a nonnative fish that was introduced into Arizona in 1921. They are native to the upper Mississippi River basin. These bass are most often bronze to brownish in color, with dark vertical bars on the sides.
In contrast to the largemouth bass, the upper jaw does not extend beyond the rear margin of
the eye. The eye is reddish in color and there is a shallow notch in the dorsal fin. The soft
dorsal fin has 13 to 15 rays. Length can vary between 12 and 22 inches, and smallmouth
bass can weigh between 8 ounces and 7 pounds. [5]
Location and Habitat: Within its native range the smallmouth bass seems most abundant in pools of streams that
consist of a substantial proportion of riffle habitat, clean, rocky, hard bottoms, and gradients
of 0.5 to about 5.0 m per km. In large rivers and lakes, smallmouth bass tend to congregate
over hard, stony bottoms, where currents are present. At the present time, smallmouth bass occur in the mainstream of the Colorado River, in the
Verde River system, and throughout the Salt River Basin below about 2,200 meters in
elevation. [6]
Temperature: Temperatures may be the most important single factor limiting distribution
of smallmouth bass. Faster
growth rates of adult smallmouth bass are generally associated with higher
summer temperatures. Faster growth rates
occur in southern reservoirs, resulting in earlier death than in northern
regions. In the summer, bass inhabit warmer shoreline areas of
large lakes in the North and deeper, cooler waters in the South. Growth does not begin until water
temperatures reach 10-14° C. Field data indicate that adults prefer temperatures of about 21-27° C
in the summer. Smallmouth bass have been reported "sunning"
themselves in pools with water temperatures of about 26.7° C in summer. [7]
Temperature preferences of smallmouth bass vary considerably depending on
the acclimation temperature.
Smallmouth bass acclimated at 2.2-30.0° C selected temperatures of 20-32° C in
laboratory tests. Adult bass in the laboratory preferred
temperatures of 28° C to 31° C. Optimum growth rates in the lab occurred at temperatures
from 26-29° C. Upper lethal
temperatures for adults were above 32.3° C. [8]
When temperatures drop to 15-20° C, adults seek deep, dark areas. At about 10° C, bass become
inactive and seek shelter. At 6-7° C, most smallmouth bass are beneath
the rock substrate, with few remaining on top. The lower
lethal temperature is near freezing. Bass will congregate around
warm springs in winter when available.[9]
Turbidity and Dissolved Oxygen: Smallmouth bass apparently can tolerate periodic turbidity,
although excessive turbidity and siltation will reduce a population. Hubert and Lackey (1980) reported a typical smallmouth
bass habitat to have very low turbidity, usually < 25 JTU, and almost never
> 75 JTU (except under flood conditions when turbidity is sometimes as high as
250 JTU).[10]
Salinity: Smallmouth occur at pH levels of 5.7 to 9,
although optimum pH is 7.9-8.1. Butler (1972) found
that smallmouth bass cover-seeking behavior was reduced at pH levels < 6, and
the lower lethal pH level was 3. Smallmouth bass populations are more productive in clearer, less fertile
reservoirs several years after impoundment that have low total dissolved
solids (TDS = 100-350 ppm). [11]
Reproduction: Smallmouth bass spawn in spring, usually mid-April to July, depending on
geographical location and water temperature. Cleary (1956) observed a 45 day
post-nesting period for smallmouth spawning in Iowa streams.
Smallmouth bass spawn on rocky lake shoals, river shallows, or backwaters
or move into creeks or tributaries to- spawn. The species requires a clean
stone, rock, or gravel substrate for spawning.
Studies show that the habitat condition during spawning is the most important
factor for year class strength in smallmouth bass.
Nest building and spawning occur when the water temperature is
12.8-21.0° C, but most activity occurs at or above 15° C. [12]
Mature females may contain 2000-15,000 golden yellow eggs. Males may spawn with several females on a single nest. On average each nest contains about 2,500 eggs, but nests may contain as many as 10,000 eggs. Eggs hatch in about 10 days if water temperatures are in the mid-50's (°F), but can hatch in 2-3 days if temperatures are in the mid-70's (°F). Males guard the nest from the time eggs are laid until fry begin to disperse, a period of up to a month. [13]
Bass spawning videos
Food: The diet of smallmouth bass changes from small to large food items as the
fish grow. Fry feed on microcrustaceans. Juvenile smallmouth bass eat larger insects, crayfish, and fish. Adults primarily feed on fish and crayfish in both lakes and
streams. The diet is influenced by abundance and availability of prey. [14]
Risk: In Arizona, smallmouth bass reportedly are responsible for eliminating or reducing some
populations of native fishes. [15]
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Spike flows
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Large fluctuations in water level can affect reproductive success
(Pflieger 1975; Montgomery et al . 1980). Ideal spawning conditions include
one or more substantial rises in water level a week or two prior to bass
nesting (Pflieger 1975) and relatively stable water levels while nesting is in
progress (Watson 1955; Pflieger 1975). Rising water may flush nest areas with
cold water, causing nest desertion and halting embryo development (Watt 1959;
Montgomery et al. 1980). Falling water levels may drive guarding males off,
limit water circulation around eggs, and increase predation, resulting in
lower reproductive success (Neves 1975; Montgomery et al. 1980).[16]
Fry seem to be especially vulnerable to flood conditions and fluctuating
water levels (Larimore 1975). A rapid drop in water level may trap them in
areas where they will become dessicated (Montgomery et al . 1980). A stream
rise of only a few inches may displace advanced fry newly risen from the nest
(Webster 1954). Most fry remain in shallow water (Doan 1940; Forney 1972),
although some may be found at depths of 4.6.-6.1 m (Stone et al. 1954; Forney
1972). Fry 20-25 mm in length cannot maintain themselves in current velocities
> 200 mm/sec (Larimore and Duever 1968). An increase in turbulence during
flood conditions creates conditions with which smallmouth fry appear unable to
cope (Webster 1954). Fry cannot tolerate and are displaced at high turbidities
(2,000 JTU) combined with an increase in water velocity, but they will not be
displaced at moderate turbidities (250 JTU) (Larimore 1975). Low water temperatures during flood conditions will reduce fry swimming ability (Larimore
and Duever 1968).[17]
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