Difference between revisions of "Smallmouth Bass Page"
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| − | [[ | + | As anticipated, the 2024 Cool Mix flows negatively impacted smallmouth bass in Glen Canyon and Grand Canyon, leading to reduced growth, a decline in relative abundance, and no confirmed recruitment or spawning observed. [[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|(1)]] |
| − | + | [[File:2023_2025SMBcatches.png|center|300px]] [https://www.usbr.gov/uc/progact/amp/twg/2025-07-10-twg-meeting/uc-gcdamp-twg-smallmouthbassnonnatives-508-20250710.pdf https://www.usbr.gov/uc/progact/amp/twg/2025-07-10-twg-meeting/uc-gcdamp-twg-smallmouthbassnonnatives-508-20250710.pdf] | |
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| − | + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;"> How many smallmouth bass are being collected in Grand Canyon? </h2> | |
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| − | + | In 2023, the mean daily catch per unit effort (CPUE) of smallmouth bass between GCD and Lees Ferry increased from May to July. The highest CPUE in 2023 occurred in October when young of the year smallmouth bass grew to catchable sizes. In 2024, SMB relative abundance remained stable throughout the year, indicating that no new fish were growing to catchable size and suggesting that no reproduction occurred. Both juvenile and large adults (i.e., >200 mm), were more abundant in 2023 compared to 2024. Due to low capture rates in 2023-2024, relative abundance estimates for smallmouth bass below Lees Ferry were not calculated. | |
| − | + | Length-frequency data revealed the presence of a new year class of smallmouth bass in 2023, shown by the larger proportion and abundance of juvenile smallmouth bass (20–30 mm) in June-August and growing larger into the fall. In 2024, we did not see fish within this size class (20-30 mm) supporting a lack of bass reproduction below the dam in 2024. [[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|(2)]] | |
| − | + | {| class="sortable wikitable" style="text-align:center;" | |
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| + | !Year | ||
| + | !Adult smallmouth bass | ||
| + | !Juvenile smallmouth bass | ||
| + | !Total | ||
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| + | |2026||1||70||71 (August AMWG) | ||
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| + | |2025||5||46||[https://www.usbr.gov/uc/progact/amp/amwg/2026-02-26-amwg-meeting/uc-gcdamp-amwg-smallmouthbass-508-20260226.pdf 51] | ||
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| + | |2024||2||250||[[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx| 254]] | ||
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| + | |2023||1||1,112||[[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|1,113]] | ||
| + | |- | ||
| + | |2022||2||360||[[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|362]] | ||
| + | |- | ||
| + | |2003-2021||11||11||[[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|22]] | ||
| + | |} | ||
| − | + | Difficulty arises when trying to compare the number of smallmouth bass being caught from year to year due to changes in capture methodologies. For example, if one were to focus removal efforts in an area where smallmouth bass are concentrated like in a backwater or a slough, the catch per unit effort (or CPUE) would be higher that the CPUE of normal monitoring which by design is done at sites that are selected more at random. | |
| + | Additionally, the level of effort can change from year to year and those changes may not be reflected when presenting the number of fish being caught. For example, if one caught a certain number of fish one year using a certain level of effort and then the next year they doubled the level of effort but only caught a third more fish, the CPUE (and relative abundance) would be smaller the second year even though more fish were caught. Both capture methodologies and level of effort has changed from year to year with the numbers of smallmouth bass listed above. | ||
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| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;"> Where are smallmouth bass currently being collected in Grand Canyon? </h2> | ||
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| + | Most smallmouth bass continue to be collected in the first couple of miles below Glen Canyon Dam. | ||
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| + | From 2022-2024, the furthest downstream smallmouth bass were captured was at RM 16.38R in November 2023. The furthest downstream adult smallmouth bass (>200mm) was captured within the slough at RM -12.19L in July 2023. Distribution of smallmouth bass in 2024 was similar to that observed in 2023 with the exception that bass were largely absent from the -12 mi slough. A total of 252 smallmouth bass were captured across all projects, with 191 of those fish captured in the Glen Canyon reach (RM –15.71 to RM 0). Bass captures in the PBR reach (RM 0 to RM 8) increased slightly in 2024 (13 individuals captured vs 8 in 2023), suggesting that large-scale migration downstream is not yet occurring. [[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|(3)]] | ||
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| + | [[File:2024smbCapturelocations.jpg|center|300px]] [https://www.usbr.gov/uc/progact/amp/twg/2024-10-30-twg-meeting/20241030-SmallmouthBassUpdates-508-AMWQD.pdf https://www.usbr.gov/uc/progact/amp/twg/2024-10-30-twg-meeting/20241030-SmallmouthBassUpdates-508-AMWQD.pdf] | ||
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| + | In 2024, environmental DNA (eDNA) samples from both sides of the river every mile from RM 0 - RM 20 and every 5 miles from RM 25 – RM 65. This spans the distance from Lees Ferry to the Little Colorado River. Positive eDNA detections were made down to RM 37.5. In large rivers, eDNA can be detected more than 100 km downstream though the strongest signals are typically within 10–100 meters of the source [https://onlinelibrary.wiley.com/doi/10.1111/1755-0998.13934]. | ||
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| + | [[File:2024_smb_edna.png|center|300px]] [https://www.usbr.gov/uc/progact/amp/amwg/2025-08-21-amwg-meeting/uc-gcdamp-amwg-smallmouthbass-508-20250821.pdf https://www.usbr.gov/uc/progact/amp/amwg/2025-08-21-amwg-meeting/uc-gcdamp-amwg-smallmouthbass-508-20250821.pdf] | ||
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| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;"> Where are they coming from? Entrainment or local reproduction below the dam? </h2> | ||
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| + | Maintaining Lake Powell elevations above 3590 ft would likely minimize entrainment and create downstream conditions that minimize smallmouth bass population growth. Entrainment increases dramatically as reservoir elevation declines below 3530 ft (see the [https://gcdamp.com/index.php/Entrainment Entrainment page]). [https://gcdamp.com/images_gcdamp_com/d/d4/Eppehimer_etal_2025_Declining_res_elevations_non-native_fish_passage_CJFAS.pdf] | ||
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| + | There appears to be two cohorts of juvenile bass being collected in the Glen Canyon Dam tailwater. One from 2022 and another from 2023 (2025 Annual Reporting Meeting). Young of the year smallmouth bass were collected from the [https://gcdamp.com/index.php/The_-12_Mile_Slough -12 Mile Slough] in both 2022 and 2023 indicating that local reproduction occurred in the slough in those two years. The catch distribution of smallmouth bass in the tailwater being greater nearest the dam indicates either entrainment and/or additional spawning right below the dam may have been also contributing. | ||
| + | [[File:2022_2023CohortGrowth.png|200px]] [https://www.usbr.gov/uc/progact/amp/twg/2025-04-10-twg-meeting/uc-gcdamp-twg-arm-smallmouthbass-coolmix-508-20250410.pdf https://www.usbr.gov/uc/progact/amp/twg/2025-04-10-twg-meeting/uc-gcdamp-twg-arm-smallmouthbass-coolmix-508-20250410.pdf] | ||
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| + | Release temperatures from Glen Canyon Dam suggesting that epilimnetic releases were made in 2022 (yellow) and 2023 (purple), but not in 2021 (orange) or 2024 (green). Release temperatures for 2024 are presented here without mixing the cooler bypass releases. A higher likelihood of entrainment is associated with epilimnetic releases like those that occurred in 2022 and 2023. | ||
| + | [[File:2021_2024releasetempertures.png|center|300px]] | ||
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| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;"> How long have smallmouth bass been found in Grand Canyon? </h2> | ||
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| + | Smallmouth bass have been sporadically captured in Grand Canyon since 2003. Captures continue to be concentrated below Glen Canyon Dam but individuals have been caught in the past at the Little Colorado River confluence and in western Grand Canyon near the Lake Mead inflow. They were first introduced into Lake Powell in 1982 [https://wayneswords.net/threads/smallmouth-bass-history-in-lake-powell.1340/] and there are populations in ponds and lakes in the upper Little Colorado River and were first detected in Lake Mead in 2000 [https://www.usbr.gov/uc/progact/amp/twg/2024-01-25-twg-meeting/20240125-AnnualReportingMeeting-SmallmouthBassPopulationModelingImplications-508-UCRO.pdf]. | ||
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| + | [[File: SmallmouthCaptures.png|center|300px]] [https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-Presentation-508-UCRO.pdf https://www.usbr.gov/uc/progact/amp/twg/2022-04-13-twg-meeting/20220413-Near-TermThreatSmallmouthBassEstablishmentGlenCanyonDam-Presentation-508-UCRO.pdf] | ||
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| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;"> What is the likelihood of smallmouth bass establishment below Glen Canyon Dam and what risk might that pose to humpback chub in Grand Canyon? </h2> | ||
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| + | =='''[https://doi.org/10.1093/tafafs/vnaf027 William E Pine, Josh Korman, Richard A Valdez, Carl J Walters. 2025. Assessing the potential for Smallmouth Bass population establishment in Grand Canyon. Transactions of the American Fisheries Society, Volume 154, Issue 5, September 2025, Pages 469–489]'''== | ||
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| + | The expansion of nonnative Smallmouth Bass (Micropterus dolomieu) into the Grand Canyon ecosystem downstream of Glen Canyon Dam is widely seen as a potential threat to native fishes, particularly the Humpback Chub (Gila cypha), which is listed as threatened under the Endangered Species Act. This concern stems from observations in the Green and upper Colorado rivers, where dam-modified habitats and reservoir introductions have allowed Smallmouth Bass to become established and impact native fish species. Our objective was to compare habitat conditions between a Smallmouth Bass population center in the Green River— where Humpback Chub are rare— and two major Humpback Chub population centers in the Colorado River in Grand Canyon, where Smallmouth Bass invasion is a conservation concern, to help assess invasion risk and inform management actions downstream of Glen Canyon Dam. | ||
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| + | We developed a conceptual model of Smallmouth Bass recruitment potential emphasizing the roles of temperature, turbidity, and the timing of these conditions, based on literature for Smallmouth Bass and Largemouth Bass (Micropterus nigricans). Using U.S. Geological Survey gauge station data, we summarized daily temperature and turbidity patterns near a known Smallmouth Bass population center in the middle Green River (near Jensen, Utah) and compared those with conditions in the Colorado River in Glen Canyon and Grand Canyon downstream from Glen Canyon Dam. We then used our conceptual model of Smallmouth Bass early life history, along with observed turbidity and temperature patterns, to explore how irregular recruitment could affect Smallmouth Bass population dynamics and individual growth potential using an age-structured population model and a bioenergetics model. | ||
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| + | Although warm reservoir releases due to low water storage in Lake Powell has made temperature regimes in Glen and Grand canyons more similar to those of the middle Green River, the Colorado River ecosystem in Grand Canyon has a very different turbidity regime from that of the middle Green River. The Colorado River in Grand Canyon becomes very turbid during late summer and fall, when tributaries experience flash floods during the North American monsoon season. These periods of high turbidity coincide with critical early life stages of Smallmouth Bass, likely limiting foraging efficiency, growth, and overwinter survival. Our population model indicates that rapidly growing and sustained Smallmouth Bass populations require successful recruitment at least once every 2 years, but empirical data suggest that these conditions occur only every 5–7 years in Grand Canyon. This would make long-term establishment of self-sustaining Smallmouth Bass populations in Grand Canyon unlikely or at least highly uncertain. | ||
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| + | Despite the presence of nonnative predators, Humpback Chub populations in Grand Canyon have expanded during the past 10–15 years, very likely due to favorably warm river temperature. While Smallmouth Bass in Glen or Marble canyons (between Glen Canyon Dam and the Little Colorado River) may pose a downstream dispersal risk (and potentially a threat to Humpback Chub populations), high turbidity conditions downstream of the Little Colorado River reduce the likelihood of persistent local Smallmouth Bass recruitment. These results suggest that Smallmouth Bass invasion risk to Humpback Chub is not uniform across the Colorado River basin and should be evaluated in the context of the different site-specific water quality and flow regime conditions, other known risks to Humpback Chub, and Humpback Chub population size in different parts of the basin. | ||
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| + | =='''[https://doi.org/10.1093/tafafs/vnag026 Charles B Yackulic, Drew E Eppehimer, Lindsey A Bruckerhoff, Anna Amidon, Kevin R Bestgen, Nathaniel Bransky, Kimberly L Dibble, Maria Dzul, Lindsay E Hansen, Brian D Healy, M Tildon Jones, Chris Michaud. 2026. Comment on “Assessing the potential for Smallmouth Bass population establishment in Grand Canyon.” Transactions of the American Fisheries Society]'''== | ||
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| + | Our primary concern is that Pine et al. rely on data from a less turbid segment of the Green River to determine the turbidity tolerance of SMB in Grand Canyon; however, SMB are distributed much farther downstream in the Green River in segments with a greater number of warm, turbid days than the segment of the Green River used by Pine et al. | ||
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| + | Our second major concern is that Pine et al. do not consider contemporary or reasonable future temperature scenarios in their analysis of turbid, warm days in the Grand Canyon ecosystem. The frequency of warm, turbid days increases under the future temperature scenario. This occurs because low reservoir levels allow 16°C to be exceeded earlier in the calendar year during months when the Colorado River in the Grand Canyon ecosystem is less likely to be turbid (e.g., May and June), thereby adding a few warm, turbid days and many warm, clear days. | ||
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| + | Our third concern is that Pine et al. do not address the relatively mild winters in the Grand Canyon ecosystem, which are expected to facilitate SMB establishment in this ecosystem. | ||
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| + | Our final concern is that Pine et al. make conclusions regarding the population impacts of SMB on HBC without a thorough, empirically supported analysis based on absolute abundances of different life stages of SMB (e.g., subadults and adults) and the per capita impacts of these SMB life stages on different life stages of HBC (e.g., juveniles, subadults, and adults). | ||
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| + | =='''[https://doi.org/10.1093/tafafs/vnag027 William E Pine, Josh Korman, Richard A Valdez, Carl J Walters. 2026. Response to Comment on “Assessing the potential for Smallmouth Bass population establishment in Grand Canyon.” Transactions of the American Fisheries Society]'''== | ||
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| + | We evaluated key claims by Yackulic et al. (2026) about Smallmouth Bass risk to Humpback Chub in Marble and Grand canyons. Three conclusions emerge. | ||
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| + | First, Yackulic et al.’s (2026) empirical analysis primarily addresses turbidity frequency but does not directly evaluate the seasonal turbidity structure emphasized by Pine et al. (2025). Warm-season turbidity patterns differed consistently between the Green River and sites in Marble and Grand canyons. At the Green River site near Jensen, turbidity events occurred mainly in June, when water temperatures generally approach the Smallmouth Bass spawning threshold (Pine et al., 2025), reflecting late snowmelt runoff and showing that late-summer and early-fall turbidity events were uncommon in most years. In contrast, early summer at the Colorado River site in Marble Canyon, upstream of the Little Colorado River, and the Colorado River in Grand Canyon downstream of the Little Colorado River, was typically dominated by clear days, while late summer and fall were marked by more frequent and longer turbidity events associated with the North American Monsoon and sediment inputs from key tributaries, including the Paria and Little Colorado rivers. This seasonal difference matters because late-summer and early-fall turbidity directly overlaps the period when age-0 Smallmouth Bass must feed and grow before winter. A turbid day in June, before or near spawning, is not equivalent to a turbid day in August or September, when age-0 Smallmouth Bass are accumulating energy reserves and growing toward sizes that improve overwinter survival. | ||
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| + | Yackulic et al. (2026) argue that Grand Canyon’s relatively short winters reduce the minimum size that age-0 Smallmouth Bass must reach to survive their first winter. We agree that this mechanism is consistent with Shuter et al. (1980), who showed that overwinter survival depends on both body size at the start of winter and the duration of the starvation period. However, this argument increases establishment probability only if age-0 Smallmouth Bass first grow sufficiently during the preceding summer and fall. Winter duration and summer turbidity are not independent filters; they act sequentially on the same cohort. Shorter winters may lower the size threshold for overwinter survival, but late-season turbidity affects whether age-0 Smallmouth Bass can reach that threshold in the first place. If monsoon- driven turbidity repeatedly interrupts foraging from August through October, age-0 Smallmouth Bass may still fail to reach even the lower threshold associated with Grand Canyon’s shorter winters. | ||
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| + | Second, Green River–like Smallmouth Bass densities within the observed range of the Humpback Chub survival model produced modest estimated reductions in juvenile survival. | ||
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| + | Third, the broader history of nonnative predators in Marble and Grand canyons is consistent with the hypothesis that warm-season turbidity and other downstream habitat conditions can limit high-density populations of visually oriented predators downstream from Glen Canyon. The Little Colorado River inflow population has persisted over the past three decades despite periods of high densities of nonnative Rainbow Trout, Brown Trout, and Green Sunfish, as well as main-stem Colorado River conditions considered too cold for Humpback Chub recruitment. | ||
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| + | Several factors could reduce the likelihood that Smallmouth Bass predation alone would cause immediate population level effects across Grand Canyon. The history of several nonnative predators suggests that high abundance in Glen Canyon does not necessarily translate into high abundance in Marble or Grand canyons, where turbidity and other downstream habitat conditions may limit dispersal, survival, or persistence. Additionally, the Little Colorado River likely provides a partial refuge for Humpback Chub given its frequent high turbidity and elevated dissolved solids. Western Grand Canyon Humpback Chub aggregations occur far downstream from Glen Canyon, where Lake Powell may serve as a Smallmouth Bass source population. Finally, Smallmouth Bass were detected in western Grand Canyon before the formation of Pearce Ferry Rapid but did not develop a documented self-sustaining population there despite thermal suitability and proximity to a Lake Mead source population. These observations do not mean that Smallmouth Bass pose little risk to Humpback Chub; they indicate that risk depends on whether Smallmouth Bass can establish and maintain high densities in places and seasons where vulnerable Humpback Chub life stages are exposed. | ||
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| + | The central disagreement between Pine et al. (2025) and Yackulic et al. (2026) is not whether turbidity can influence Smallmouth Bass abundance; both papers accept that it can. Rather, the disagreement concerns which features of the turbidity regime are most relevant for recruitment risk. Yackulic et al. (2026) emphasize the frequency of warm, turbid days during the warm season. Pine et al. (2025) emphasize seasonal turbidity structure: when turbid events occur, how intense they are, and whether they persist long enough to interrupt feeding and growth during critical early life stages. The key question for Grand Canyon is therefore not whether Smallmouth Bass recruitment could ever occur but whether favorable recruitment years would occur often enough to sustain the high Smallmouth Bass densities necessary to reverse more than a decade of Humpback Chub population growth. | ||
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| − | *[https:// | + | *[https://gcdamp.com/index.php/SEIS_Bypass_Releases SEIS Bypass Releases Page] |
| − | *[https:// | + | *[https://gcdamp.com/index.php/The_-12_Mile_Slough The -12 Mile Slough Page] |
| − | *[https:// | + | *[https://gcdamp.com/index.php/Turbidity Turbidity Page] |
| + | *[https://gcdamp.com/index.php/Entrainment Entrainment Page] | ||
| + | *[http://gcdamp.com/index.php/Nonnative_Invasive_Aquatic_Species Nonnative Invasive Aquatic Species Page] | ||
*[https://www.usbr.gov/uc/progact/amp/amwg/2023-02-16-amwg-meeting/20230216-InvasiveFishSpeciesBelowGlenCanyonDam-508-UCRO.pdf 2023 Invasive Fish Species Below Glen Canyon Dam: A Strategic Plan to Prevent, Detect and Respond] | *[https://www.usbr.gov/uc/progact/amp/amwg/2023-02-16-amwg-meeting/20230216-InvasiveFishSpeciesBelowGlenCanyonDam-508-UCRO.pdf 2023 Invasive Fish Species Below Glen Canyon Dam: A Strategic Plan to Prevent, Detect and Respond] | ||
*[http://gcdamp.com/index.php?title=National_Park_Service_Expanded_Non-native_Aquatic_Species_Management_Plan_and_EA 2019 National Park Service Expanded Non-native Aquatic Species Management Plan] | *[http://gcdamp.com/index.php?title=National_Park_Service_Expanded_Non-native_Aquatic_Species_Management_Plan_and_EA 2019 National Park Service Expanded Non-native Aquatic Species Management Plan] | ||
*[http://gcdamp.com/index.php?title=2013_National_Park_Service_Comprehensive_Fisheries_Management_Plan 2013 National Park Service Comprehensive Fisheries Management Plan] | *[http://gcdamp.com/index.php?title=2013_National_Park_Service_Comprehensive_Fisheries_Management_Plan 2013 National Park Service Comprehensive Fisheries Management Plan] | ||
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*[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf Habitat suitability information: Smallmouth bass] | *[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf Habitat suitability information: Smallmouth bass] | ||
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| + | *[https://gcdamp.com/index.php/SEIS_Bypass_Releases SEIS Bypass Releases at Glen Canyon Dam] | ||
*[https://coloradoriverrecovery.org/uc/wp-content/uploads/sites/2/2022/02/TechnicalReport-nna-Bestgen-2018-SmallmouthBassStudyPlan.pdf Evaluate effects of flow spikes to disrupt reproduction of smallmouth bass in the Green River downstream of Flaming Gorge Dam] | *[https://coloradoriverrecovery.org/uc/wp-content/uploads/sites/2/2022/02/TechnicalReport-nna-Bestgen-2018-SmallmouthBassStudyPlan.pdf Evaluate effects of flow spikes to disrupt reproduction of smallmouth bass in the Green River downstream of Flaming Gorge Dam] | ||
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| + | '''2026''' | ||
| + | *[[Media:Pine et a 2025 GCsmb.pdf|Pine et al. 2025. Assessing the potential for Smallmouth Bass population establishment in Grand Canyon. Transactions of the American Fisheries Society]] | ||
| + | *[[Media:Pine et al 2025 GCsmb supplementary data.docx| Supplemental Information for Pine et al. 2025. Assessing the potential for Smallmouth Bass population establishment in Grand Canyon. Transactions of the American Fisheries Society]] | ||
| + | *[https://academic.oup.com/tafs/advance-article/doi/10.1093/tafafs/vnag026/8734007?searchresult=1 Yackulic et al. 2026. Comment on “Assessing the potential for Smallmouth Bass population establishment in Grand Canyon.” Transactions of the American Fisheries Society] | ||
| + | *[https://academic.oup.com/tafs/advance-article/doi/10.1093/tafafs/vnag027/8734929?searchresult=1 Pine et al. 2026. Response to Comment on “Assessing the potential for Smallmouth Bass population establishment in Grand Canyon.” Transactions of the American Fisheries Society] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2026-02-26-amwg-meeting/uc-gcdamp-amwg-smallmouthbass-508-20260226.pdf Smallmouth Bass and Nonnative Fish Monitoring in 2025] | ||
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| + | '''2025''' | ||
| + | *[[Media:Eppehimer_etal_2025_Declining_res_elevations_non-native_fish_passage_CJFAS.pdf|Eppehimer et al., 2025, Declining reservoir elevations following a two-decade drought increase water temperatures and non-native fish passage facilitating a downstream invasion: Canadian Journal of Fisheries and Aquatic Sciences]] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2025-12-11-twg-meeting/uc-gcdamp-twg-sbahgupdate-508-20251211.pdf SBAHG Update] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2025-12-11-twg-meeting/uc-gcdamp-twg-smallmouthbass-508-20251211.pdf Smallmouth Bass and Other High-Risk Nonnatives] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2025-08-21-amwg-meeting/uc-gcdamp-amwg-smallmouthbass-508-20250821.pdf Smallmouth Bass and other Nonnatives] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-07-10-twg-meeting/20240710-SmallmouthBassUpdates-508-AMWD.pdf Smallmouth Bass Updates] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-07-10-twg-meeting/20240710-SBAHGUpdate-508-AMWD.pdf SBAHG Update] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-07-10-twg-meeting/20240710-TuckerTrawlHydroacousticUpdate-508-AMWD.pdf Tucker Trawl and Hydroacoustic Update] | ||
| + | *[https://www.nps.gov/articles/000/smallmouth-bass-threat.htm Smallmouth Bass: Not a Small Threat to the Grand Canyon] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2025-04-10-twg-meeting/uc-gcdamp-twg-arm-smallmouthbass-coolmix-508-20250410.pdf Response of Smallmouth Bass and other species to cool mix] | ||
'''2024''' | '''2024''' | ||
| − | *[https:// | + | *[https://digitalcommons.usu.edu/cgi/viewcontent.cgi?article=1304&context=etd2023 Friesen, Barrett T., "Invasion Potential of Nonnative Fishes Through a Large Western Dam Into a Prized and Vulnerable Ecosystem" (2024). All Graduate Theses and Dissertations, Fall 2023 to Present. ] |
| − | *[https://www. | + | *[[Media:LTEMP_SEIS_Cool_Mix_Flow_Summary_Report_Draft4.docx|2024 Cool Mix Flow Summary Report]] |
| − | *[https://www.usbr.gov/ | + | *[https://qcnr.usu.edu/coloradoriver/files/research/2024-SMB-Report-3.pdf Review of Smallmouth Bass Management in the Colorado River Ecosystem] |
| − | *[https://www.usbr.gov/ | + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-10-30-twg-meeting/20241030-SBAHGUpdate-508-AMWQD.pdf SBAHG Update] |
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-10-30-twg-meeting/20241030-SmallmouthBassUpdates-508-AMWQD.pdf Smallmouth Bass Updates] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/twg/2024-10-30-twg-meeting/20241030-2024BassNestMonitoring-508-AMWQD.pdf 2024 Bass Nest Monitoring] | ||
| + | *[https://www.usbr.gov/uc/progact/amp/amwg/2024-08-22-amwg-meeting/20240822-SmallmouthBassUpdates-508-AMWQD.pdf Smallmouth Bass Updates ] | ||
*[https://www.usbr.gov/uc/progact/amp/amwg/2024-02-29-amwg-meeting/20240229-FisheriesReviewAnnualReportingFY2023-508-UCRO.pdf Fisheries Review: Annual Reporting FY2023 ] | *[https://www.usbr.gov/uc/progact/amp/amwg/2024-02-29-amwg-meeting/20240229-FisheriesReviewAnnualReportingFY2023-508-UCRO.pdf Fisheries Review: Annual Reporting FY2023 ] | ||
*[https://www.usbr.gov/uc/progact/amp/twg/2024-01-25-twg-meeting/20240125-AnnualReportingMeeting-SmallmouthBassPopulationModelingImplications-508-UCRO.pdf Smallmouth Bass Population Modeling and Implications ] | *[https://www.usbr.gov/uc/progact/amp/twg/2024-01-25-twg-meeting/20240125-AnnualReportingMeeting-SmallmouthBassPopulationModelingImplications-508-UCRO.pdf Smallmouth Bass Population Modeling and Implications ] | ||
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'''2023''' | '''2023''' | ||
*[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-TWGMeeting-SBAHGUpdateNonnativeFishStrategicPlan-508-UCRO.pdf SBAHG Update & Nonnative Fish Strategic Plan: Review and Discussion] | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-TWGMeeting-SBAHGUpdateNonnativeFishStrategicPlan-508-UCRO.pdf SBAHG Update & Nonnative Fish Strategic Plan: Review and Discussion] | ||
| − | |||
*[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-AnnualReportingMeeting-SmallmouthBassDepletionEffortGlenCanyonNRA-508-UCRO.pdf Smallmouth Bass Depletion Effort in Glen Canyon National Recreation Area ] | *[https://www.usbr.gov/uc/progact/amp/twg/2023-01-26-twg-meeting/20230126-AnnualReportingMeeting-SmallmouthBassDepletionEffortGlenCanyonNRA-508-UCRO.pdf Smallmouth Bass Depletion Effort in Glen Canyon National Recreation Area ] | ||
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Mature females may contain 2000-15,000 golden yellow eggs. Males may spawn with several females on a single nest. On average each nest contains about 2,500 eggs, but nests may contain as many as 10,000 eggs. Eggs hatch in about 10 days if water temperatures are in the mid-50's (°F), but can hatch in 2-3 days if temperatures are in the mid-70's (°F). Males guard the nest from the time eggs are laid until fry begin to disperse, a period of up to a month. [https://www.fs.usda.gov/Internet/FSE_DOCUMENTS/fsbdev3_018493.pdf] | Mature females may contain 2000-15,000 golden yellow eggs. Males may spawn with several females on a single nest. On average each nest contains about 2,500 eggs, but nests may contain as many as 10,000 eggs. Eggs hatch in about 10 days if water temperatures are in the mid-50's (°F), but can hatch in 2-3 days if temperatures are in the mid-70's (°F). Males guard the nest from the time eggs are laid until fry begin to disperse, a period of up to a month. [https://www.fs.usda.gov/Internet/FSE_DOCUMENTS/fsbdev3_018493.pdf] | ||
| − | + | '''Movement:''' | |
| − | + | A smallmouth bass movement study in the Yampa River noted that smallmouth bass can move great distances upstream and downstream (Hawkins et al. 2009). | |
| − | + | ||
'''Food: '''The diet of smallmouth bass changes from small to large food items as the | '''Food: '''The diet of smallmouth bass changes from small to large food items as the | ||
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'''Risk: '''In Arizona, smallmouth bass reportedly are responsible for eliminating or reducing some | '''Risk: '''In Arizona, smallmouth bass reportedly are responsible for eliminating or reducing some | ||
populations of native fishes. [https://www.usbr.gov/lc/phoenix/biology/azfish/pdf/smallmouthbass.pdf] | populations of native fishes. [https://www.usbr.gov/lc/phoenix/biology/azfish/pdf/smallmouthbass.pdf] | ||
| + | |||
| + | '''Bass spawning videos:''' | ||
| + | *[https://www.youtube.com/watch?v=50m8DoiKVVU Video 1] | ||
| + | *[https://www.youtube.com/watch?v=zBDDliPHdVk Video 2] | ||
| + | |||
| + | |- | ||
| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Will smallmouth bass spawn in the fall?</h2> | ||
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| + | |||
| + | [https://storre.stir.ac.uk/handle/1893/29308 Cantin (1994)] found that natural spawning occurs in spring (April to July) when the photophase (duration of daylight) is long and increasing and water temperatures are between 15°C to 25°C. Under controlled laboratory conditions, she found that spawning could be induced from January to the end of June independently of the photophase. Individual spawnings indicated that the same female can spawn more than once per season and at different times of the year if the right developmental and spawning conditions are met. At the end of the experiments, the fish left from all the groups were stocked, in mid-July, into a lake. August and September spawnings were then observed. These spawnings occurred with temperature fluctuations between 15-18°C and decreasing photoperiod. | ||
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| − | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Spike | + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">Smallmouth Bass Spike Flows</h2> |
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Fry seem to be especially vulnerable to flood conditions and fluctuating | Fry seem to be especially vulnerable to flood conditions and fluctuating | ||
water levels (Larimore 1975). A rapid drop in water level may trap them in | water levels (Larimore 1975). A rapid drop in water level may trap them in | ||
| − | areas where they will become | + | areas where they will become desiccated (Montgomery et al . 1980). A stream |
rise of only a few inches may displace advanced fry newly risen from the nest | rise of only a few inches may displace advanced fry newly risen from the nest | ||
(Webster 1954). Most fry remain in shallow water (Doan 1940; Forney 1972), | (Webster 1954). Most fry remain in shallow water (Doan 1940; Forney 1972), | ||
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and Duever 1968).[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] | and Duever 1968).[https://apps.dtic.mil/sti/pdfs/ADA323294.pdf] | ||
| + | |- | ||
| + | ! <h2 style="margin:0; background:#cedff2; font-size:120%; font-weight:bold; border:1px solid #a3b0bf; text-align:left; color:#000; padding:0.2em 0.4em;">When and how were smallmouth bass introduced to Lake Powell (by Wayne Gustaveson) [https://wayneswords.net/threads/smallmouth-bass-history-in-lake-powell.1340/] </h2> | ||
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| + | Lake Powell filled in 1980 after 17 years of constantly rising water levels. The consequences were dramatic. Largemouth bass and crappie had dominated the fishery in the filling reservoir. That changed, as the full reservoir began fluctuating up and down. Annual fluctuating water levels combined with daily wind and wave action uprooted brush in the fluctuation zone. The lake topped out at 3700 feet while lake level changed an average of 20 to 25 feet in any given year. Low water levels in the fall allowed some quick growing annual weeds to survive but perennial plants did not have enough time to become established before spring floods covered the ground once more. The shoreline now consisted of barren rock and sand instead of terrestrial desert vegetation that could be used by young bass and crappie as nursery cover when the flood covered the new brush each spring. | ||
| + | |||
| + | Monitoring the situation would not fix this problem as largemouth bass and crappie populations would crash as a result of inadequate nursery cover. The rapidly expanding striped bass population would incorrectly be blamed for the demise of largemouth bass and crappie. It was time for active management. My research indicated that smallmouth bass would reproduce and perform well in rocky substrate based on the simple principle that smallmouth fry hide in rocks when danger threatens. Largemouth bass and crappie fry must hide in brushy cover to ward off predation. Without brush, largemouth survival was poor but with two thousand miles of rocky shoreline smallmouth bass would thrive. This research was presented to the staff in Salt Lake who approved the concept. | ||
| + | |||
| + | This is where it gets a bit tricky. Our striped bass hatchery in Big Water, Utah was out of production after striped bass natural reproduction was discovered in 1979. The hatchery was not used in 1980 or 1981. But we had a hatchery and the means to grow smallmouth bass. All that was needed was approval from Salt Lake, an adult bass brood source, and the ponds under my direction could turn out smallmouth bass fry in short order. The plans were approved and we went to work. Smallmouth bass were readily available in Flaming Gorge Reservoir. In spring 1982 we grabbed our fishing rods, caught 150 bass, and relocated them from the cold northern Utah border to the Southern Utah tropics. | ||
| + | |||
| + | It takes time to understand all the nuances of raising a new species of fish in a hatchery. The first year is often a learning experience where mistakes are made that lead to correct operations in the coming years. It was no different in 1982. We had not used the hatchery for two years and we had never raised smallmouth bass before, so the learning curve was steep. We made some mistakes but managed to produce about 10,000 bass fry when 10 times that many were expected. | ||
| + | |||
| + | My instructions from Salt Lake were to produce as many bass as possible. The highest priority was to place the first year’s progeny in Starvation Reservoir in Northern Utah. Second priority was to stock another lake in the north and third priority, if we had enough fish, would be Lake Powell. | ||
| + | |||
| + | Obviously with our meager production results there were not enough bass to stock all three lakes. The decision was made to place all the production from 1982 in Starvation Reservoir. The ponds were lowered and the two to three inch fingerlings were harvested over a three day period. It took a long time to harvest fish from our antiquated ponds that did not have catch basins. We had to lower the ponds individually and seine each pond a number of times to collect fish, which were then placed in a small 10x10 ft holding net until all the ponds were drained. On the third day the ponds were all harvested with all smallmouth fry placed in one holding net in one of the ponds. The stocking airplane was scheduled to arrive the next morning. | ||
| + | |||
| + | The plane landed at sunrise and we were dipping small bass out of the large holding net in the pond. Bass fry were dipped from the big net and placed in seven small holding nets. Then the small nets were suspended in a fish tank and rushed a half-mile up the hill to the flat road where the plane had landed. There were just enough fish in each net to fill one of the seven small, wet fish compartments on the plane. These 10,000 fish were extremely crowded in the plane’s holding tank. Time was critical to get them loaded and stocked as soon as possible. The flight took over two hours from Wahweap to Starvation so we worked fast and hard to make it happen. The plane was airborne again in less than an hour. | ||
| + | |||
| + | After the intense and focused effort to load fish in a timely manner it was time to relax and drain the fishpond. As the water went down it became obvious that the pond contained more than a few escapees. I looked at the net and saw that overnight maybe 500 but less than 1000 bass had exited our holding net out of one small hole. The plane was gone and it was not possible to put these fish on that flight. There were not enough fish to justify another expensive cross state flight to Starvation or to another lake in northern Utah. We had a few extra smallmouth bass. | ||
| + | |||
| + | In 1982 there were no cell phones, no Internet, and at the hatchery there was no telephone. We did not even have a hatchery building. We had a hatchery stocking truck and 500 extra fish. Someone needed to make a decision. My first option was to send the fish to Starvation. That goal had been met. The second option was to bring the plane back to put 500 fish in another lake. That was not economically feasible. Option three was to put 500 fish in the stocking truck and drop them off in Lake Powell on the way home. We could meet our third priority and take care of priority two next year when more fish would be raised. | ||
| + | |||
| + | It made sense, and it was economically practical. I stocked smallmouth bass in Lake Powell by driving the hatchery truck down Crosby Canyon road, opening the tube, and letting the fish flow down the pipe. It only took an hour and no extra funds. It was the right thing to do and I didn’t give it an extra thought. | ||
| + | |||
| + | In those days we had much more autonomy. We had to make decisions based on our directions and understanding. We communicated with our superiors by phone and by writing a monthly report, which was sent by US mail. My smallmouth stocking was completed before having access to the phone so I just included that incident in my monthly report and sent the letter. About three days later when the letter arrived in Salt Lake my phone rang shortly after the letter was opened. | ||
| + | |||
| + | The conversation was short. Don Andriano asked, “You did what?” I responded “Yes, I stocked 500 smallmouth bass in Lake Powell as it was the best choice of the options available to me at the time.” | ||
| + | |||
| + | Don asked, “Did you know that we did not have permission to do that?” I responded “No I was never told that, only that Lake Powell was third priority for stocking smallmouth bass this year.” | ||
| + | |||
| + | What was done was final. The conversation was over. I am sure Don had some explaining to do before the Colorado River Fish and Wildlife Council. In retrospect, asking forgiveness may have been a better choice than asking permission, which likely would never have been granted as I later learned with my proposal to introduce rainbow smelt into Lake Powell. | ||
| + | |||
| + | One result of my management action was satisfaction in knowing that smallmouth bass predictably performed magnificently along the rocky shoreline of the lake just as my research indicated they would. Anglers have caught millions of smallmouth bass over the past 30 years. | ||
| + | |||
| + | Another consequence was the threat of smallmouth bass predation on native species in the river running through the Grand Canyon. Fortunately, smallmouth did not populate the river as it was too cold for successful bass reproduction. Bass and native chubs and suckers need water temperatures above 60 degrees to successfully reproduce. The cold tailwater coming out of Lake Powell (46F) does not often provide adequate spawning temperature so bass and native fish do not reproduce in the mainstem Colorado River in most years. | ||
| + | |||
| + | Largemouth and smallmouth bass have been good teammates in Lake Powell. As predicted, largemouth bass numbers crashed shortly after the lake filled. The lake over filled in 1983 covering brush along the shoreline once more. That gave largemouth an extra year and the population boomed until the lake stabilized and gradually declined from 1984 to 1992. When the brush was gone largemouth bass numbers plummeted. | ||
| + | |||
| + | Drought-induced low lake levels in the early 90s allowed more brush to grow on the shoreline. Rising water in 1993 and 1994 covered the new brush and largemouth bass responded immediately as fry produced found nursery cover and survived at a high rate. That population peak lasted from 1994 to 1996 as a generation of bass completed their life cycle. The lake then stabilized again eliminating brush in the fluctuation zone. The next drought occurred at the same time as the Y2K scare that had folks wondering if computers would melt down as the new century was ushered in. From 2000 to 2004 Lake Powell steadily declined to a new low of 3555 MSL. New brush grew along the wet shoreline, as lake bottom became sandy beach once more. When the lake came back up from 2005 to 2008, largemouth bass again peaked in numbers and regained the high population level last seen when the lake filled and overfilled in 1981 to 1984. | ||
| + | |||
| + | I have included this technical discussion and graphics to show that largemouth bass respond to lake level and subsequent brush growth or elimination. Management of largemouth bass in Lake Powell would be possible if lake level could be controlled and regulated to allow new brush growth on demand. If there was a large red button on my desk that allowed me to control lake level, I could produce a consistent trophy-sized supply of largemouth bass and crappie at Lake Powell just by manipulating brush cover. | ||
| + | Some have logically stated that striped bass predation has been responsible for the demise of largemouth in Lake Powell. The data indicates that the recent largemouth population boom from 2007 to 2011 occurred when the striped bass population was also abundantly strong. That seems impossible if striped bass predation was responsible for a decline in largemouth bass abundance. There is not a direct correlation or relationship between striped bass and largemouth populations. They are often separated without much contact by their affinity for different habitat types. Striped bass live in open water and prefer to feed on shad while largemouth live in shallow brushy habitat and feed on shad, sunfish and crayfish. | ||
Latest revision as of 20:44, 18 August 2026
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Smallmouth Bass (Micropterus dolomieui)The predatory threat of invasive and large-bodied piscivorous taxa such as smallmouth bass in the upper Colorado River basin is substantial. For example, based on results of a bioenergetics model, Johnson et al. (2008) ranked smallmouth bass as the most problematic invasive species because of their high abundance, habitat use that overlaps with most native fishes, and ability to consume a wide variety of life stages of native fishes (Bestgen et al. 2008). Expanded populations of piscivores such as smallmouth bass are a major impediment to conservation actions aimed at recovery efforts for the four endangered fishes in the upper Colorado River basin (U.S. Fish and Wildlife Service 2002a, b, c, d). [1] |
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